By Amcarrig
Basic Anatomy
Tridacnid clams have shells
consisting of two halves, or valves, which are joined at the top. Muscles and
ligaments on each side of the shell hold the shell closed and pull the shell
open. Species of tridacnid clams can be easily identified by the shape,
symmetry and texture of their shells.
All clams have a
tissue structure called a mantle. One function of the mantle is to increase the
surface area of the clam so that it receives maximum exposure to the light. It
also precipitates calcium carbonate which forms the clam's shell. The mantle
contains symbiotic algae (zooxanthellae) and light sensory organs which
contribute to a clam's coloration and pattern. The zooxanthellae produce
photosynthesis-derived food for its host while the light sensory organs serve to
position the clam for maximum light exposure, to protect the clam against
excessive light and UV radiation and to warn the clam of potential predators.
The
mantle is whole except for where it is broken by two siphons; the inhalant
siphon, which is often fringed with fine tentacles that strain larger particles
out of the water, and the exhalent siphon which is tube like.
Photo of T.
crocea showing the mantle, the inhalant siphon, the exhalent siphon and the
gills (photo courtesy of A. Carriglio)
Clams are filter
feeders. Water and food are drawn in through the inhalant siphon to the gills
where food is caught. The gills also draw oxygen from the water. From the
gills, the food is transported along a groove to an organ which pushes it into
the clam's mouth. The exhalent siphon carries away the water. Clams get some
of their food from filtering the water, however, they get a lot more of their
nourishment from their symbiotic algae.
The byssal gland of a clam secretes
filaments called byssal threads which hold the clam to the substrate. Because
the larger clams, such as T. gigas, T. derasa, and H. hippopus get so large,
they will eventually lose the gland and rely on their size and weight to hold
them in place. T maxima and T. crocea are rock boring clams, therefore, their
byssal glands excrete a substance that dissolves the substrate to which they are
attached which helps them to burrow into it.
Lighting requirements
Some tridacnid clams require moderate to high lighting
and some require intense lighting. Those that require moderate to high lighting
are T. squamosa, T. derasa, H hippopus and T. gigas. Those that require intense
lighting are T. maxima and T. crocea. If lighting in the aquarium is not
sufficient, the clam may display higher than normal mantle extension. This
usually means that the clam is trying to extend its mantle for maximum light
exposure because of insufficient lighting.
A guide to
buying and caring for tridacnid clams
Make sure that you have enough lighting
and room in your aquarium to support the clam long term.
Only purchase clams that respond quickly
to shadow, touch or other stimulus by closing its shell.
The mantle should not have any tears or
other damage to it. The mantle should extend past the edge of the shell (H.
hippopus being the exception).
Check for signs of gaping (shell is open
wider than what is considered normal, little or no mantle extension, intake
siphon open wider than what is considered normal, slow closing response to
stimulus). A gaping clam should not be purchased.
You should not see any torn or loose
tissue hanging from the bottom of the clam. Some byssal threads may be visible,
but no solid tissue should be hanging.
Place clams on the appropriate substrate
and away from any corals or other inverts that may sting it. Do not place the
clam where it is constantly blasted by strong and direct water currents.
Because the exhalent siphons of a clam are
capable of quickly expelling large amounts of water, you'll want keep this in
mind if you decide to place your clam high up in the aquarium, close to your
lighting system.
If you need to move a clam to another
location in your aquarium, it is very important that you not tug at or pull on
the clam if it is attached to the substrate. It's best to cut the byssal
threads as close to the substrate as possible with a razor blade until the clam
is no longer attached and then relocate it. One way to avoid having the clam
attach to something that cannot be moved or attach in a place where you cannot
reach the byssal threads, is to place a flat piece of live rock or a shell
underneath the clam. Once the clam attaches itself to this piece of rock or
shell, it can be easily moved. The rock/shell also prevents predation of the
byssal muscle from below the sandbed.
Certain angelfish, triggerfish, puffers,
butterfly fish, wrasses and other fish have been know to eat clams as do certain
crabs and shrimp. Therefore, spontaneous fish or invert purchases should be
avoided if you are keeping clams or plan to keep them in the future.
Many clams can harbor parasitic boring
(pyramid) snails which can wreak havoc in a tank full of tridacnid clams. Look
for rice grain-sized, cream colored spots near the base or in the scutes of the
clam, or, at night, along the upper edge of the shell. If the clam is attached
to a rock, check by lifting the clam a short distance off the rock and look
underneath. You are looking for small (0.08-0.2 inch / 2-5 mm long) snails.
Remove all of these snails. If you have a quarantine tank, quarantine the clam
until you are sure that all of the snails are removed. Also check for small,
jelly like egg masses and remove them as well. Don't confuse the jelly-like
mass some clams excrete around their byssus opening for these egg
masses
When introducing the clam to your
aquarium, it's common for your fish and shrimp to check it out for any desirable
food that may have come in on the clam's shell. This should only be a concern
if the animal begins to bite and/or tear at the mantle.
The number one cause of a clam's
demise is usually water quality.
Calcium is the main building
block for clams and should be present in the water at levels of at least 280
mg/L for growth to occur. More rapid, natural growth is seen when calcium is in
the range of 400-480 mg/L.
Strontium is incorporated in the
shell along with calcium and should also be provided for optimum growth. The
addition of iodide to the aquarium will also enhance growth and color in
clams.
High pH and temperatures can
cause problems. Do not let the aquarium exceed 82 degrees or a pH above 8.3.
Maintain a dkh of 7.9.
Too high or low a salinity can
cause the death of a clam. Try to keep specific gravity between 1.023 and
1.025.
|
TRIDACNID
IDENTIFICATION GUIDE
|
|
Type |
Mantle
Color/Characteristics |
Shell |
Approx. Adult
Size |
Byssal |
Siphon |
Lighting
Requirement |
Placement |
Feeding |
|
T. Maxima
|
Blue, purple,
gray, brown, and yellow are common.
Black specimens exist but are somewhat rare. Spotted, striped, or blotched. Patches of
solid color are more common than multi-colored patterns. |
Asymmetrical.
5 very distinct ribs. Moderately defined, tightly spaced scutes. |
16" |
Medium to large byssal
opening.
Semi-rock boring
|
Small, simple tentacles on
inhalant |
Intense |
Preferably on rockwork. Brightly colored
specimens require brighter lighting than brown ones. Brown T. maximas should be placed lower in
the aquarium to prevent them from getting shocked by strong
lighting |
Phytoplankton only when clam is
3" or smaller. Light and supplemental
phyto when
larger. |
|
T. Crocea
|
Blue,
purple, yellow, green, gold, orange, and brown.
Lines, spots, or blotches of yellow, blue, and green can also be
seen. |
Asymmetrical
5 to 6 moderately defined
ribs. Scutes
are tightly spaced but not well defined. |
6" |
Large byssal opening. Rock boring. |
Small, simple tentacles on
inhalant |
Intense |
Preferably on rockwork. Clams with bright
coloring (especially blue) should be placed high in the
aquarium |
Phytoplankton only when clam is
3" or smaller. Light and supplemental
phyto when
larger. |
|
T. Derasa
|
Wavy
striped patterns or spots. Can be found in black, white, blue, yellow, and
orange color combinations, but most mantles have a golden color. Some have
bright green or blue lines. |
Symmetrical. . 5
to 7 moderately defined ribs that lack scutes. |
20" |
Narrow/small byssal
opening. |
Large complex tentacles on inhalant |
Strong |
On rockwork or on
sand |
Phytoplankton only when clam is
3" or smaller. Light and supplemental
phyto when
larger. |
|
T.
Squamosa
|
Beige, brown, or gold. Some can have blue or green blotches
and some have stripes running parallel to the
shell. |
Symmetrical
4-5 large, very defined ribs.
Large, sharply defined, widely spaced scutes. |
16" |
Medium to non-existent byssal
opening. |
Large complex tentacles on inhalant |
Strong |
On rockwork or on sand
|
Phytoplankton only when clam is
3" or smaller. Light and supplemental
phyto when
larger. |
|
T. Gigas
|
Blue, green,
golden-brown, or yellow in color. Iridescent spots may also be present,
especially near the edge of the mantle |
Asymmetrical
4 to 5 defined ribs. Juveniles have
some scutes on their shells but adults do
not. |
Up to
4' |
Small to nonexistent byssal
opening. |
No tentacles on
inhalant |
Strong |
On
sand |
Phytoplankton only when clam is
3" or smaller. Light and supplemental
phyto when
larger. |
|
H.
hippopus
|
Brown-green with some deep
yellow striping. The mantle of H.
hippopus does not extend past the shell edge as do the other
tridacnids |
Asymmetrical and very
distinct.
Typically has 7 to 8 well defined ribs which lack scutes. |
14" |
Small byssal
opening. |
No tentacles in
inhalant |
Strong |
On
sand |
Phytoplankton only when clam is
3" or smaller. Light and supplemental
phyto when
larger. |
References
Identifying
the Giant Clams, James W. Fatherree, M. Sc., Reefkeeping On-line Magazine. http://www.reefkeeping.com/issues/2002-07/jf/feature/index.php
Fish & Chips, A Monthly
Marine Newsletter, Elizabeth M. Lukan 8/17/99.
http://animal-world.com/encyclo/fishnchips/aug99/fnc0899.htm
Giant Clams: A Comprehensive
Guide to the Identification and Care of Tridacnid Clams, Daniel Knop, Ricordea
Publishing, July 1996.